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On both of my Macs, since updating to 10.4.3, dashboard and expose keep forgetting the function keys that are assigned to them. This happens sometimes even whilst running, i.e. not after a sleep or restart.
I've experienced this with Dashboard, which occasionally forgets F12. It seems to affect only keyboard shortcuts, as my Exposé screen corners are never forgotten.
Very, very annoying. This started happening for me before 10.4.3, btw.
I've had this happen several times, in both 10.4.2 and 10.4.3. Specifically, my keyboard and mouse shortcuts for Exposé, Dashboard, and the dictionary all disappear. I have filed a bug report on it.